The railway line between Lewes and Uckfield has been closed since 1969. While there have been discussions and studies regarding its reinstatement, no further work is currently planned due to the lack of an economic case, according to East Sussex County Council. The line was part of the broader Wealden Line.
This article published by the Friends of Lewes is a very useful guide to the history of railway lines in the Lewes area going back to the original 19th C construction of the railway network: CLICK HERE
Here is the route of the Lewes to Uckfield line as depicted in the 20o3 OS map:


Above copyright OS Maps 2003
Larger versions on these maps are available here: